Botany - systematization, evolution, paleobotany, floristics, morphology, botanical geography, biosystematization

Graduated from USSR subtropical cultures Institute of Tbilisi (1932). Dr. of Biological Sciences (1943). Prof. (1944), acad. (1941, corr. member 1943)

Chief of the department of the plants of Insitute of botany by name V.Komarova Academy of Science of Russian Federation (1963), adviser of the director (since 1986)

Full member АН the Russian Federation, АН the USA, Linevsky societies of London, German academy of naturalists by name Leopoldin, the Polish and Norwegian academies

Winner Gos. Premiums of the USSR. Awarded with Labour Red Banner, Friendship of peoples awards, medals "For valorous work" (2) and "Gold" medals of NAS RA (2005). Hero of Sots. Work. Laureate of V.Komarova and R.Alertona awards

Chief of Faculty of morphology and systematization of plants (1938-1948), director of Institute botany NAS RA (1944), Prof. (1949-1961), dean of the Biological fact (1951-1954) of St.-Petersburg University, director (1977-1986) of Insitute of botany by name V.Komarova Academy of Science of Russian Federation. Distinguished scientist of Armenia (1967)