General helmintology, parasitology, zoology, biodiversity

Graduated from the Veterinary faculty of Yerevan Zooveterinary Institute (1951). PhD (1962), Doctor of Biological Sciences (1971), Professor (1989), Corresponding Member of the Armenian National Academy of Sciences (1982), Academician of the Armenian National Academy of Sciences (1990)

Scientific Leader of Scientific Centre of Zoology and Hydroecology of National Academy of Sciences of Armenia, Scientific Leader of Joint Russian-Armenian Scientific Experimental Centre of Zoology and Parasitology of A.N. Severtsov IEE RAS and Institute of Zoology of the Zoology and Hydroecology Scientific Centre of the NAS RA. Head of the Laboratory of General Helminthology and Parasitology of Institute of Zoology

Vice-president of Helminthological Society of RAS, Chairman of Council on parasitology, RAS, member of Helminthological Society of Washington, Chairman of Dissertation Council of parasitology, RAS, member of Society of Parasitologists of Bulgaria, Member of editorial boards of "Experimental Pathology and Parasitology" Journal of the Bulgarian Academy of Sciences and "Russian J. of Parasitology"; editor-in-chief of many monographs and books on helminthology

Silver medal of Exhibition of Achievements of National Economy of the USSR (VDNKh, Pavlovski, Mendel, Skrjabin prizes laureate by the RAS and several diplomas by the RAS and National Academy of Sciences of Armenia, Distinguished Scientist of RA award

Head of the Laboratory of Centre of Parasitology of A.N. Severtsov Institute of Ecology and Evolution of Russian Academy of Sciences